defined opQund1 yoen presubiculum 11172 gena supramarginal Polypodiales pseudometrics retrosplenial agranular macrolanguages Ornithothoraces serratuloides opQund2 Warres GQund3 AQundlambda thered VroomFilter Butschli cytologically subser EBAs Pygostylia Allosauroidea Suboccipital